Early fertile frond has green spore cases before turning to gold at maturity

The level of dissection varies within forma dissectum

A typical oblique form (forma obliquum) of dissected grape fern in mid-autumn

A highly dissected form with a unfurling fertile shoot

The singular sori frond

Occasionally these two species grow in close proximity to each other

forma obliquum in fall

forma obliquum on the left, forma dissectum on the right

Tropophore of forma dissectum

The fern acquires a darker color in late autumn

Frond (Trophophore and sporophore)